:root {
  color-scheme: dark;
  --ink: #f4f7ff;
  --ink-soft: #c4cde0;
  --ink-muted: #8c98b0;
  --night: #050812;
  --night-raised: #0a1020;
  --panel: rgba(16, 24, 44, 0.82);
  --panel-solid: #10182c;
  --panel-hover: #15213b;
  --line: rgba(164, 184, 224, 0.18);
  --line-strong: rgba(164, 184, 224, 0.35);
  --cyan: #66e3ff;
  --cyan-soft: rgba(102, 227, 255, 0.13);
  --violet: #a995ff;
  --violet-soft: rgba(169, 149, 255, 0.14);
  --amber: #ffd479;
  --amber-soft: rgba(255, 212, 121, 0.13);
  --green: #79e6bd;
  --green-soft: rgba(121, 230, 189, 0.12);
  --red: #ff9b9b;
  --red-soft: rgba(255, 155, 155, 0.12);
  --shadow: 0 24px 80px rgba(0, 0, 0, 0.32);
  --radius-sm: 0.55rem;
  --radius-md: 0.9rem;
  --radius-lg: 1.35rem;
  --content: 76rem;
  --reading: 48rem;
  --font-sans: Inter, ui-sans-serif, system-ui, -apple-system, BlinkMacSystemFont, "Segoe UI",
    sans-serif;
  --font-mono: "SFMono-Regular", Consolas, "Liberation Mono", monospace;
}

*,
*::before,
*::after {
  box-sizing: border-box;
}

html {
  scroll-behavior: smooth;
  scroll-padding-top: 5.5rem;
}

body {
  min-width: 18rem;
  margin: 0;
  background:
    radial-gradient(circle at 12% -8%, rgba(87, 103, 255, 0.19), transparent 32rem),
    radial-gradient(circle at 92% 12%, rgba(45, 207, 236, 0.1), transparent 27rem),
    linear-gradient(180deg, #060a15 0%, var(--night) 52%, #070b14 100%);
  color: var(--ink);
  font-family: var(--font-sans);
  font-size: 1rem;
  line-height: 1.62;
  text-rendering: optimizeLegibility;
}

body::before {
  position: fixed;
  z-index: -1;
  inset: 0;
  background-image:
    radial-gradient(circle, rgba(255, 255, 255, 0.56) 0 1px, transparent 1.4px),
    radial-gradient(circle, rgba(113, 217, 255, 0.32) 0 1px, transparent 1.4px);
  background-position:
    0 0,
    4.8rem 7rem;
  background-size:
    11rem 11rem,
    17rem 17rem;
  content: "";
  mask-image: linear-gradient(to bottom, black, transparent 64rem);
  opacity: 0.14;
  pointer-events: none;
}

img,
svg {
  display: block;
  max-width: 100%;
}

button,
input,
select {
  font: inherit;
}

button,
select {
  cursor: pointer;
}

a {
  color: var(--cyan);
  text-underline-offset: 0.2em;
}

a:hover {
  color: #b0f1ff;
}

:focus-visible {
  outline: 3px solid var(--amber);
  outline-offset: 3px;
}

::selection {
  background: rgba(102, 227, 255, 0.26);
  color: #fff;
}

[hidden] {
  display: none !important;
}

.skip-link {
  position: fixed;
  z-index: 100;
  top: 0.75rem;
  left: 0.75rem;
  padding: 0.65rem 0.9rem;
  border-radius: var(--radius-sm);
  background: var(--ink);
  color: var(--night);
  font-weight: 800;
  transform: translateY(-180%);
  transition: transform 160ms ease;
}

.skip-link:focus {
  color: var(--night);
  transform: translateY(0);
}

.site-header__inner,
main > section,
main > article,
.site-footer__inner {
  width: min(calc(100% - 2rem), var(--content));
  margin-inline: auto;
}

.site-header {
  position: sticky;
  z-index: 30;
  top: 0;
  border-bottom: 1px solid var(--line);
  background: rgba(5, 8, 18, 0.83);
  backdrop-filter: blur(18px) saturate(1.25);
}

.site-header__inner {
  display: flex;
  min-height: 4.5rem;
  align-items: center;
  justify-content: space-between;
  gap: 1.5rem;
}

.brand {
  display: inline-flex;
  align-items: center;
  gap: 0.7rem;
  color: var(--ink);
  font-size: 0.9rem;
  font-weight: 850;
  letter-spacing: 0.08em;
  text-decoration: none;
  text-transform: uppercase;
}

.brand::before {
  width: 1.05rem;
  height: 1.05rem;
  border: 2px solid var(--cyan);
  border-radius: 50%;
  box-shadow:
    0 0 0 0.28rem rgba(102, 227, 255, 0.08),
    0 0 1.2rem rgba(102, 227, 255, 0.35);
  content: "";
}

.brand strong {
  color: var(--cyan);
}

.site-nav {
  display: flex;
  align-items: center;
  gap: 0.35rem;
}

.site-nav a {
  padding: 0.45rem 0.7rem;
  border-radius: 999px;
  color: var(--ink-soft);
  font-size: 0.88rem;
  font-weight: 650;
  text-decoration: none;
}

.site-nav a:hover {
  background: var(--cyan-soft);
  color: var(--ink);
}

main {
  display: block;
  overflow: clip;
}

main > section {
  padding-block: clamp(3.8rem, 7vw, 6.5rem);
}

main > section + section {
  border-top: 1px solid rgba(164, 184, 224, 0.1);
}

.eyebrow {
  margin: 0 0 0.75rem;
  color: var(--cyan);
  font-family: var(--font-mono);
  font-size: 0.76rem;
  font-weight: 750;
  letter-spacing: 0.14em;
  text-transform: uppercase;
}

h1,
h2,
h3,
h4 {
  color: var(--ink);
  font-weight: 780;
  letter-spacing: -0.035em;
  line-height: 1.08;
}

h1 {
  max-width: 15ch;
  margin: 0;
  font-size: clamp(3rem, 8vw, 6.8rem);
}

h2 {
  margin: 0 0 1rem;
  font-size: clamp(2rem, 4.8vw, 3.8rem);
}

h3 {
  margin: 0;
  font-size: 1.2rem;
}

h4 {
  margin: 0;
  font-size: 0.88rem;
  letter-spacing: -0.01em;
}

.button,
.pagination button {
  display: inline-flex;
  min-height: 2.75rem;
  align-items: center;
  justify-content: center;
  padding: 0.58rem 0.95rem;
  border: 1px solid var(--line-strong);
  border-radius: var(--radius-sm);
  background: rgba(164, 184, 224, 0.07);
  color: var(--ink);
  font-size: 0.86rem;
  font-weight: 750;
  text-decoration: none;
  transition:
    border-color 160ms ease,
    background-color 160ms ease,
    transform 160ms ease;
}

.button:hover,
.pagination button:hover:not(:disabled) {
  border-color: var(--cyan);
  background: var(--cyan-soft);
  color: var(--ink);
  transform: translateY(-1px);
}

.button--primary {
  border-color: var(--cyan);
  background: var(--cyan);
  color: #031018;
}

.button--primary:hover {
  background: #a5f1ff;
  color: #031018;
}

/* Hero -------------------------------------------------------------------- */

.hero {
  position: relative;
  isolation: isolate;
  display: grid;
  min-height: min(43rem, calc(100svh - 4.5rem));
  grid-template-columns: minmax(0, 1.45fr) minmax(18rem, 0.55fr);
  align-content: center;
  column-gap: clamp(2rem, 7vw, 7rem);
  padding-block: clamp(5rem, 10vw, 9rem);
}

.hero::after {
  position: absolute;
  z-index: -1;
  top: 8%;
  right: -8rem;
  width: min(54vw, 38rem);
  aspect-ratio: 1;
  border: 1px solid rgba(102, 227, 255, 0.18);
  border-radius: 50%;
  background:
    radial-gradient(circle at 38% 34%, rgba(255, 255, 255, 0.9) 0 0.16%, transparent 0.4%),
    radial-gradient(circle at 45% 45%, rgba(102, 227, 255, 0.18), rgba(71, 60, 185, 0.05) 38%, transparent 66%);
  box-shadow:
    inset 0 0 4rem rgba(102, 227, 255, 0.05),
    0 0 8rem rgba(58, 89, 212, 0.08);
  content: "";
  opacity: 0.85;
}

.hero__content {
  max-width: 52rem;
  align-self: center;
}

.hero h1 em {
  color: transparent;
  background: linear-gradient(100deg, var(--cyan), #b8b0ff 70%);
  background-clip: text;
  font-family: ui-serif, Georgia, serif;
  font-weight: 520;
}

.hero__lede {
  max-width: 45rem;
  margin: 1.35rem 0 0;
  color: var(--ink-soft);
  font-size: clamp(1.05rem, 2vw, 1.28rem);
}

.hero-actions,
.study-actions {
  display: flex;
  flex-wrap: wrap;
  gap: 0.75rem;
  margin-top: 1.75rem;
}

.last-checked {
  display: flex;
  flex-wrap: wrap;
  align-items: center;
  gap: 0.65rem 1.25rem;
  margin: 1rem 0 0;
  color: var(--ink-muted);
  font-family: var(--font-mono);
  font-size: 0.74rem;
}

.status-label {
  display: inline-flex;
  align-items: center;
  gap: 0.5rem;
}

.status-label::before {
  width: 0.55rem;
  height: 0.55rem;
  border-radius: 50%;
  background: var(--green);
  box-shadow: 0 0 0.8rem rgba(121, 230, 189, 0.6);
  content: "";
}

.hero-biology {
  position: relative;
  width: min(100%, 25rem);
  aspect-ratio: 1;
  align-self: center;
  justify-self: center;
}

.cell-ring {
  position: absolute;
  top: 50%;
  left: 50%;
  border: 1px solid rgba(102, 227, 255, 0.25);
  border-radius: 46% 54% 52% 48% / 56% 44% 56% 44%;
  transform: translate(-50%, -50%) rotate(-12deg);
}

.cell-ring--one {
  width: 35%;
  height: 32%;
  border-color: rgba(169, 149, 255, 0.4);
}

.cell-ring--two {
  width: 64%;
  height: 59%;
  transform: translate(-50%, -50%) rotate(18deg);
}

.cell-ring--three {
  width: 94%;
  height: 88%;
  border-style: dashed;
  transform: translate(-50%, -50%) rotate(-6deg);
}

.cell-nucleus {
  position: absolute;
  top: 50%;
  left: 50%;
  width: 5rem;
  height: 5rem;
  border: 1px solid rgba(169, 149, 255, 0.58);
  border-radius: 46% 54% 49% 51%;
  background:
    radial-gradient(circle at 38% 34%, rgba(255, 255, 255, 0.92) 0 2%, transparent 3%),
    radial-gradient(circle, rgba(169, 149, 255, 0.8), rgba(92, 67, 185, 0.46) 58%, transparent 70%);
  box-shadow:
    0 0 0 0.7rem rgba(169, 149, 255, 0.05),
    0 0 3rem rgba(102, 227, 255, 0.2);
  transform: translate(-50%, -50%);
}

.cell-particle {
  position: absolute;
  width: 0.78rem;
  height: 0.78rem;
  border-radius: 50%;
  background: var(--amber);
  box-shadow:
    0 0 0 0.32rem rgba(255, 212, 121, 0.08),
    0 0 1.2rem rgba(255, 212, 121, 0.68);
}

.cell-particle--one {
  top: 20%;
  left: 34%;
}

.cell-particle--two {
  top: 62%;
  left: 79%;
  width: 0.58rem;
  height: 0.58rem;
  background: var(--cyan);
  box-shadow: 0 0 1rem rgba(102, 227, 255, 0.8);
}

.cell-particle--three {
  top: 76%;
  left: 21%;
  width: 0.46rem;
  height: 0.46rem;
  background: var(--green);
  box-shadow: 0 0 1rem rgba(121, 230, 189, 0.75);
}

.metrics-grid {
  display: grid;
  grid-column: 1 / -1;
  grid-template-columns: repeat(4, minmax(0, 1fr));
  gap: 1px;
  margin: 2.5rem 0 0;
  overflow: hidden;
  border: 1px solid var(--line);
  border-radius: var(--radius-lg);
  background: var(--line);
}

.metric {
  display: flex;
  min-height: 8.5rem;
  flex-direction: column;
  padding: 1.25rem;
  background: var(--panel-solid);
}

.metric__value {
  order: -1;
  display: block;
  margin: 0;
  color: var(--ink);
  font-family: var(--font-mono);
  font-size: clamp(1.75rem, 3.2vw, 2.7rem);
  font-weight: 700;
  letter-spacing: -0.06em;
  line-height: 1;
}

.metric__label {
  display: block;
  margin-top: 0.75rem;
  color: var(--ink-muted);
  font-size: 0.84rem;
  line-height: 1.35;
}

/* Snapshot and discovery -------------------------------------------------- */

.snapshot-panel {
  position: relative;
  padding-block: 0;
  overflow: hidden;
  border: 1px solid var(--line);
  border-radius: var(--radius-lg);
  background:
    linear-gradient(135deg, rgba(102, 227, 255, 0.08), transparent 42%),
    var(--panel);
  box-shadow: var(--shadow);
}

.snapshot-panel::before {
  position: absolute;
  top: 0;
  bottom: 0;
  left: 0;
  width: 0.25rem;
  background: linear-gradient(var(--cyan), var(--violet));
  content: "";
}

.snapshot-panel__header {
  display: grid;
  grid-template-columns: minmax(0, 1.3fr) minmax(22rem, 0.7fr);
  align-items: center;
  gap: 2rem;
  padding: clamp(1.5rem, 4vw, 2.5rem);
}

.snapshot-panel__header h2 {
  font-size: clamp(1.85rem, 4vw, 3.2rem);
}

.snapshot-panel__header > div > p:last-child {
  max-width: 49rem;
  margin: 0.8rem 0 0;
  color: var(--ink-soft);
}

.snapshot-stats {
  display: grid;
  grid-template-columns: repeat(2, minmax(0, 1fr));
  margin: 0;
  overflow: hidden;
  border: 1px solid var(--line);
  border-radius: var(--radius-md);
  background: rgba(5, 8, 18, 0.4);
}

.snapshot-stats > div {
  min-width: 0;
  padding: 1rem;
  border-right: 1px solid var(--line);
  border-bottom: 1px solid var(--line);
}

.snapshot-stats > div:nth-child(2n) {
  border-right: 0;
}

.snapshot-stats > div:nth-last-child(-n + 2) {
  border-bottom: 0;
}

.snapshot-stats dt {
  color: var(--ink-muted);
  font-size: 0.68rem;
  letter-spacing: 0.05em;
  text-transform: uppercase;
}

.snapshot-stats dd {
  margin: 0.35rem 0 0;
  color: var(--ink);
  font-family: var(--font-mono);
  font-size: 0.88rem;
  font-weight: 700;
}

.latest-strip {
  display: grid;
  grid-template-columns: repeat(3, minmax(0, 1fr));
  gap: 1px;
  border-top: 1px solid var(--line);
  background: var(--line);
}

.latest-card {
  display: flex;
  min-width: 0;
  min-height: 10rem;
  flex-direction: column;
  padding: 1.25rem;
  background: rgba(7, 12, 25, 0.84);
  color: var(--ink);
  text-decoration: none;
  transition: background-color 160ms ease;
}

.latest-card:hover {
  background: var(--panel-hover);
  color: var(--ink);
}

.latest-card__meta {
  color: var(--cyan);
  font-family: var(--font-mono);
  font-size: 0.67rem;
  letter-spacing: 0.04em;
  text-transform: uppercase;
}

.latest-card strong {
  display: -webkit-box;
  margin-top: 0.7rem;
  overflow: hidden;
  font-size: 0.94rem;
  line-height: 1.4;
  -webkit-box-orient: vertical;
  -webkit-line-clamp: 3;
}

.latest-card small {
  margin-top: auto;
  padding-top: 0.8rem;
  overflow: hidden;
  color: var(--ink-muted);
  font-size: 0.73rem;
  text-overflow: ellipsis;
  white-space: nowrap;
}

.section-heading {
  display: flex;
  align-items: end;
  justify-content: space-between;
  gap: 2rem;
}

.section-heading > p {
  max-width: 31rem;
  margin: 0 0 0.5rem;
  color: var(--ink-soft);
}

.collections-grid {
  display: grid;
  grid-template-columns: repeat(4, minmax(0, 1fr));
  gap: 1rem;
  margin-top: 2.5rem;
}

.collection-card {
  position: relative;
  display: flex;
  min-height: 13rem;
  flex-direction: column;
  justify-content: space-between;
  gap: 1.5rem;
  padding: 1.25rem;
  overflow: hidden;
  border: 1px solid var(--line);
  border-radius: var(--radius-md);
  background: linear-gradient(155deg, rgba(23, 34, 61, 0.9), rgba(10, 16, 32, 0.92));
  color: var(--ink);
  text-decoration: none;
  transition:
    border-color 180ms ease,
    transform 180ms ease,
    background-color 180ms ease;
}

.collection-card::after {
  position: absolute;
  right: 1.15rem;
  bottom: 1.15rem;
  color: var(--cyan);
  content: "Explore →";
  font-family: var(--font-mono);
  font-size: 0.7rem;
  letter-spacing: 0.04em;
}

.collection-card:hover {
  border-color: rgba(102, 227, 255, 0.5);
  background: var(--panel-hover);
  color: var(--ink);
  transform: translateY(-3px);
}

.collection-card__count {
  color: var(--cyan);
  font-family: var(--font-mono);
  font-size: 1.65rem;
  font-weight: 700;
}

.collection-card small {
  display: block;
  margin-bottom: 0.45rem;
  color: var(--ink-muted);
  font-family: var(--font-mono);
  font-size: 0.64rem;
  letter-spacing: 0.08em;
  text-transform: uppercase;
}

.collection-card strong {
  display: block;
  max-width: 15rem;
  padding-right: 0.4rem;
  font-size: 1.06rem;
  line-height: 1.3;
}

.collection-card > span:last-child > span {
  display: block;
  margin-top: 0.55rem;
  padding-right: 0.5rem;
  padding-bottom: 1.35rem;
  color: var(--ink-muted);
  font-size: 0.79rem;
  line-height: 1.45;
}

/* Timeline ---------------------------------------------------------------- */

.timeline {
  display: flex;
  height: 18rem;
  align-items: stretch;
  gap: 0.48rem;
  margin-top: 2.5rem;
  padding: clamp(1rem, 3vw, 2rem);
  overflow-x: auto;
  border: 1px solid var(--line);
  border-radius: var(--radius-lg);
  background:
    linear-gradient(180deg, rgba(102, 227, 255, 0.035), transparent 50%),
    rgba(7, 12, 25, 0.68);
}

.timeline-bar {
  display: grid;
  min-width: 1.85rem;
  flex: 1 0 1.85rem;
  grid-template-rows: 1.4rem 1fr 2.2rem;
  align-items: end;
  color: var(--ink-muted);
  font-family: var(--font-mono);
  font-size: 0.6rem;
  text-align: center;
  text-decoration: none;
}

.timeline-bar span {
  color: var(--ink-soft);
  opacity: 0;
  transition: opacity 150ms ease;
}

.timeline-bar i {
  width: 100%;
  height: var(--bar-height);
  min-height: 3px;
  border-radius: 0.25rem 0.25rem 0 0;
  background: linear-gradient(180deg, var(--cyan), var(--violet));
  opacity: 0.78;
  transition:
    opacity 150ms ease,
    transform 150ms ease;
}

.timeline-bar small {
  align-self: start;
  writing-mode: vertical-rl;
}

.timeline-bar:hover span,
.timeline-bar:focus-visible span {
  opacity: 1;
}

.timeline-bar:hover i,
.timeline-bar:focus-visible i {
  opacity: 1;
  transform: scaleX(1.18);
}

.timeline-note {
  margin: 0.75rem 0 0;
  color: var(--ink-muted);
  font-size: 0.72rem;
  text-align: right;
}

/* Explorer ---------------------------------------------------------------- */

.explorer-shell {
  margin-top: 2.5rem;
  overflow: hidden;
  border: 1px solid var(--line);
  border-radius: var(--radius-lg);
  background: rgba(7, 12, 25, 0.62);
  box-shadow: var(--shadow);
}

.explorer-controls {
  display: grid;
  grid-template-columns: minmax(18rem, 1fr) minmax(9rem, 0.28fr) minmax(10rem, 0.28fr);
  gap: 0.8rem;
  padding: 1.25rem;
  border-bottom: 1px solid var(--line);
  background: rgba(16, 24, 44, 0.72);
}

.explorer-controls label {
  display: grid;
  gap: 0.35rem;
  min-width: 0;
  color: var(--ink-muted);
  font-size: 0.7rem;
  font-weight: 700;
  letter-spacing: 0.06em;
  text-transform: uppercase;
}

.explorer-controls input,
.explorer-controls select {
  width: 100%;
  min-width: 0;
  min-height: 2.9rem;
  padding: 0.58rem 0.72rem;
  border: 1px solid var(--line-strong);
  border-radius: var(--radius-sm);
  background: #0b1325;
  color: var(--ink);
  font-size: 0.87rem;
  letter-spacing: normal;
  text-transform: none;
}

.explorer-controls input::placeholder {
  color: #727f98;
}

.explorer-controls input:hover,
.explorer-controls select:hover {
  border-color: rgba(102, 227, 255, 0.52);
}

.explorer-controls input:focus,
.explorer-controls select:focus {
  border-color: var(--cyan);
}

.active-filters {
  display: flex;
  flex-wrap: wrap;
  gap: 0.45rem;
  padding-inline: 1.25rem;
  background: rgba(16, 24, 44, 0.52);
}

.active-filters:not(:empty) {
  padding-block: 0 1rem;
  border-bottom: 1px solid var(--line);
}

.active-filter {
  display: inline-flex;
  min-height: 1.85rem;
  align-items: center;
  gap: 0.5rem;
  padding: 0.24rem 0.55rem;
  border: 1px solid rgba(102, 227, 255, 0.28);
  border-radius: 999px;
  background: var(--cyan-soft);
  color: #baf4ff;
  font-size: 0.69rem;
}

.active-filter:hover {
  border-color: var(--cyan);
}

.active-filter span:last-child {
  font-size: 1rem;
  line-height: 0.7;
}

.filter-toggle {
  display: none;
}

.catalog-layout {
  display: grid;
  grid-template-columns: 15.5rem minmax(0, 1fr);
  align-items: start;
}

.facets {
  position: sticky;
  top: 5.5rem;
  max-height: calc(100vh - 6.5rem);
  padding: 1.25rem;
  overflow-y: auto;
  border-right: 1px solid var(--line);
  background: rgba(7, 12, 25, 0.58);
  scrollbar-color: var(--line-strong) transparent;
}

.facets__heading {
  display: flex;
  align-items: flex-start;
  justify-content: space-between;
  gap: 0.6rem;
  padding-bottom: 0.85rem;
}

.facets__heading .eyebrow {
  margin-bottom: 0.25rem;
}

.facets__heading h3 {
  font-size: 1.05rem;
}

.text-button {
  padding: 0;
  border: 0;
  background: none;
  color: var(--cyan);
  font-size: 0.72rem;
}

.text-button:hover {
  color: #b0f1ff;
  text-decoration: underline;
  text-underline-offset: 0.2em;
}

.facet-group {
  padding: 1rem 0;
  border-top: 1px solid var(--line);
}

.facet-group__heading {
  display: flex;
  align-items: center;
  justify-content: space-between;
  gap: 0.6rem;
  margin-bottom: 0.55rem;
}

.facet-group__heading span {
  color: var(--ink-muted);
  font-family: var(--font-mono);
  font-size: 0.6rem;
}

.facet-group ul {
  margin: 0;
  padding: 0;
  list-style: none;
}

.facet-group li + li {
  margin-top: 0.08rem;
}

.facet-group label {
  display: grid;
  grid-template-columns: 1rem minmax(0, 1fr) auto;
  align-items: center;
  gap: 0.48rem;
  min-height: 1.8rem;
  padding: 0.15rem 0;
  color: var(--ink-soft);
  cursor: pointer;
  font-size: 0.76rem;
}

.facet-group input {
  width: 0.92rem;
  height: 0.92rem;
  margin: 0;
  accent-color: var(--cyan);
}

.facet-group input:disabled + .facet-value {
  opacity: 0.42;
}

.facet-value {
  overflow: hidden;
  text-overflow: ellipsis;
  white-space: nowrap;
}

.facet-count {
  color: var(--ink-muted);
  font-family: var(--font-mono);
  font-size: 0.63rem;
}

.facet-more {
  margin-top: 0.45rem;
}

.catalog {
  min-width: 0;
  padding: 1.25rem;
}

.catalog-meta {
  display: flex;
  min-height: 2.1rem;
  align-items: center;
  justify-content: space-between;
  gap: 1rem;
}

.catalog-status {
  margin: 0 0 1rem;
  color: var(--ink-muted);
  font-family: var(--font-mono);
  font-size: 0.72rem;
}

.catalog-status:focus {
  outline: none;
}

.catalog-message {
  display: flex;
  min-height: 8rem;
  align-items: center;
  justify-content: center;
  gap: 0.75rem;
  color: var(--ink-muted);
  text-align: center;
}

.catalog-error {
  margin-bottom: 1rem;
  min-height: auto;
  justify-content: flex-start;
  padding: 1rem;
  border: 1px solid rgba(255, 155, 155, 0.35);
  border-radius: var(--radius-sm);
  background: var(--red-soft);
  color: #ffd3d3;
  text-align: left;
}

.loading-mark {
  width: 1.1rem;
  height: 1.1rem;
  border: 2px solid var(--line-strong);
  border-top-color: var(--cyan);
  border-radius: 50%;
  animation: spin 0.8s linear infinite;
}

.study-grid {
  display: grid;
  grid-template-columns: repeat(2, minmax(0, 1fr));
  gap: 0.85rem;
}

.study-card {
  position: relative;
  display: flex;
  min-width: 0;
  min-height: 20.5rem;
  flex-direction: column;
  padding: 1.15rem;
  border: 1px solid var(--line);
  border-radius: var(--radius-md);
  background:
    linear-gradient(150deg, rgba(22, 33, 59, 0.88), rgba(9, 15, 30, 0.92));
  transition:
    border-color 160ms ease,
    transform 160ms ease,
    background-color 160ms ease;
}

.study-card::before {
  position: absolute;
  top: -1px;
  right: 1.1rem;
  left: 1.1rem;
  height: 1px;
  background: linear-gradient(90deg, transparent, rgba(102, 227, 255, 0.5), transparent);
  content: "";
  opacity: 0;
  transition: opacity 160ms ease;
}

.study-card:hover {
  border-color: rgba(102, 227, 255, 0.45);
  background: var(--panel-hover);
  transform: translateY(-2px);
}

.study-card:hover::before {
  opacity: 1;
}

.study-card__header {
  display: flex;
  align-items: center;
  justify-content: space-between;
  gap: 0.8rem;
}

.study-card__accession,
.study-card__date {
  color: var(--ink-muted);
  font-family: var(--font-mono);
  font-size: 0.64rem;
  letter-spacing: 0.04em;
  text-transform: uppercase;
}

.study-card__accession {
  color: var(--cyan);
}

.study-card__date {
  overflow: hidden;
  text-overflow: ellipsis;
  white-space: nowrap;
}

.study-card h3 {
  margin-top: 0.75rem;
  font-size: 1.04rem;
  line-height: 1.35;
}

.study-card h3 a {
  color: var(--ink);
  text-decoration: none;
}

.study-card h3 a::after {
  position: absolute;
  inset: 0;
  content: "";
}

.study-card h3 a:hover {
  color: var(--cyan);
}

.study-card__description {
  display: -webkit-box;
  margin: 0.7rem 0 0;
  overflow: hidden;
  color: var(--ink-soft);
  font-size: 0.79rem;
  line-height: 1.55;
  -webkit-box-orient: vertical;
  -webkit-line-clamp: 4;
}

.study-card__tags {
  position: relative;
  z-index: 1;
  display: flex;
  flex-wrap: wrap;
  gap: 0.36rem;
  margin-top: 1rem;
}

.tag {
  display: inline-flex;
  max-width: 100%;
  min-height: 1.6rem;
  align-items: center;
  padding: 0.2rem 0.48rem;
  overflow: hidden;
  border: 1px solid rgba(102, 227, 255, 0.28);
  border-radius: 999px;
  background: var(--cyan-soft);
  color: #aff1ff;
  font-family: var(--font-mono);
  font-size: 0.61rem;
  font-weight: 700;
  letter-spacing: 0.02em;
  line-height: 1.2;
  text-decoration: none;
  text-overflow: ellipsis;
}

.tag--violet {
  border-color: rgba(169, 149, 255, 0.35);
  background: var(--violet-soft);
  color: #c9bfff;
}

.tag--amber {
  border-color: rgba(255, 212, 121, 0.35);
  background: var(--amber-soft);
  color: var(--amber);
}

.tag--green {
  border-color: rgba(121, 230, 189, 0.35);
  background: var(--green-soft);
  color: var(--green);
}

a.tag:hover {
  border-color: currentColor;
  color: var(--ink);
}

.study-card__footer {
  display: flex;
  align-items: center;
  justify-content: space-between;
  gap: 0.75rem;
  margin-top: auto;
  padding-top: 1rem;
  color: var(--ink-muted);
  font-size: 0.69rem;
}

.study-card__footer > span:first-child {
  overflow: hidden;
  text-overflow: ellipsis;
  white-space: nowrap;
}

.study-card__open {
  flex: 0 0 auto;
  color: var(--cyan);
  font-family: var(--font-mono);
}

.catalog-empty {
  grid-column: 1 / -1;
  min-height: 18rem;
  padding: 3.5rem 1.5rem;
  border: 1px dashed var(--line-strong);
  border-radius: var(--radius-md);
  color: var(--ink-muted);
  text-align: center;
}

.catalog-empty strong {
  color: var(--ink);
  font-size: 1.2rem;
}

.catalog-empty p {
  margin: 0.5rem 0 1.25rem;
}

.pagination {
  display: flex;
  flex-wrap: wrap;
  justify-content: center;
  gap: 0.4rem;
  margin-top: 2rem;
}

.pagination button {
  min-width: 2.65rem;
}

.pagination button[aria-current="page"] {
  border-color: var(--cyan);
  background: var(--cyan);
  color: #031018;
}

.pagination button:disabled {
  cursor: not-allowed;
  opacity: 0.38;
}

.pagination-ellipsis {
  display: inline-grid;
  min-width: 2rem;
  place-items: center;
  color: var(--ink-muted);
}

.filter-count {
  display: inline-grid;
  min-width: 1.45rem;
  height: 1.45rem;
  margin-left: 0.35rem;
  border-radius: 999px;
  background: var(--cyan);
  color: #031018;
  font-family: var(--font-mono);
  font-size: 0.68rem;
  font-weight: 800;
  place-items: center;
}

/* Methodology and footer -------------------------------------------------- */

.methodology-section {
  display: grid;
  grid-template-columns: minmax(0, 1fr) minmax(24rem, 1fr);
  align-items: center;
  gap: clamp(2rem, 7vw, 6rem);
}

.methodology-copy > p:last-child {
  max-width: 40rem;
  color: var(--ink-soft);
}

.methodology-grid {
  display: grid;
  grid-template-columns: repeat(2, minmax(0, 1fr));
  gap: 0.75rem;
  margin: 0;
}

.methodology-grid > div {
  padding: 1rem;
  border: 1px solid var(--line);
  border-radius: var(--radius-sm);
  background: var(--panel);
}

.methodology-grid dd {
  margin: 0.65rem 0 0;
  color: var(--ink-soft);
  font-size: 0.82rem;
}

.method-badge {
  display: inline-flex;
  min-height: 1.55rem;
  align-items: center;
  padding: 0.17rem 0.45rem;
  border: 1px solid var(--line-strong);
  border-radius: 0.38rem;
  font-family: var(--font-mono);
  font-size: 0.6rem;
  font-weight: 800;
  letter-spacing: 0.04em;
  text-transform: uppercase;
}

.method-badge--source {
  border-color: rgba(121, 230, 189, 0.45);
  background: var(--green-soft);
  color: var(--green);
}

.method-badge--mirror {
  border-style: double;
  border-width: 3px;
  border-color: rgba(255, 212, 121, 0.5);
  background: var(--amber-soft);
  color: var(--amber);
}

.method-badge--search {
  border-style: dashed;
  border-color: rgba(169, 149, 255, 0.55);
  background: var(--violet-soft);
  color: #c7baff;
}

.method-badge--verify {
  border-color: rgba(102, 227, 255, 0.5);
  background: var(--cyan-soft);
  color: var(--cyan);
}

.site-footer {
  margin-top: 2rem;
  border-top: 1px solid var(--line);
  background: rgba(3, 6, 13, 0.45);
}

.site-footer__inner {
  display: grid;
  grid-template-columns: minmax(0, 2fr) minmax(12rem, 1fr);
  gap: 2rem;
  padding-block: 2.5rem;
  color: var(--ink-muted);
  font-size: 0.82rem;
}

.site-footer strong {
  color: var(--ink-soft);
}

.site-footer p {
  max-width: 42rem;
  margin: 0.4rem 0 0;
}

.site-footer nav {
  display: flex;
  flex-direction: column;
  align-items: flex-start;
  gap: 0.45rem;
}

/* Study detail pages ------------------------------------------------------ */

.study-page {
  padding-block: 2.2rem 6rem;
}

.back-link {
  display: inline-flex;
  align-items: center;
  margin-bottom: 2rem;
  color: var(--ink-muted);
  font-family: var(--font-mono);
  font-size: 0.72rem;
  text-decoration: none;
}

.back-link:hover {
  color: var(--cyan);
}

.study-hero {
  display: grid;
  grid-template-columns: minmax(0, 1.35fr) minmax(18rem, 0.65fr);
  align-items: center;
  gap: clamp(2rem, 6vw, 5rem);
  min-height: 32rem;
  padding: clamp(2rem, 5vw, 4rem);
  overflow: hidden;
  border: 1px solid var(--line);
  border-radius: var(--radius-lg);
  background:
    radial-gradient(circle at 88% 12%, rgba(102, 227, 255, 0.13), transparent 25rem),
    linear-gradient(145deg, rgba(19, 29, 53, 0.96), rgba(7, 12, 25, 0.94));
  box-shadow: var(--shadow);
}

.study-hero h1 {
  max-width: 20ch;
  font-size: clamp(2.35rem, 5.7vw, 5rem);
  overflow-wrap: anywhere;
}

.study-hero__meta {
  display: flex;
  flex-wrap: wrap;
  gap: 0.55rem 1rem;
  margin-top: 1.25rem;
  color: var(--ink-muted);
  font-family: var(--font-mono);
  font-size: 0.7rem;
}

.study-hero__meta span + span::before {
  margin-right: 1rem;
  color: var(--line-strong);
  content: "·";
}

.study-hero__visual {
  position: relative;
  display: grid;
  min-height: 20rem;
  overflow: hidden;
  border: 1px solid var(--line);
  border-radius: var(--radius-lg);
  background:
    radial-gradient(circle at 50% 45%, rgba(169, 149, 255, 0.2), transparent 45%),
    rgba(5, 8, 18, 0.62);
  place-items: center;
}

.study-hero__visual::after {
  position: absolute;
  inset: 0;
  border-radius: inherit;
  box-shadow: inset 0 0 4rem rgba(5, 8, 18, 0.35);
  content: "";
  pointer-events: none;
}

.study-hero__visual img {
  width: 100%;
  height: 100%;
  min-height: 20rem;
  object-fit: cover;
}

.study-placeholder {
  position: relative;
  width: 12rem;
  aspect-ratio: 1;
  border: 1px solid rgba(102, 227, 255, 0.35);
  border-radius: 50%;
}

.study-placeholder span {
  position: absolute;
  inset: 18%;
  border: 1px solid rgba(169, 149, 255, 0.4);
  border-radius: 44% 56% 52% 48%;
}

.study-placeholder span:nth-child(2) {
  inset: 35%;
  background: rgba(169, 149, 255, 0.22);
}

.study-placeholder span:nth-child(3) {
  top: 18%;
  left: 76%;
  width: 0.8rem;
  height: 0.8rem;
  border: 0;
  border-radius: 50%;
  background: var(--amber);
  box-shadow: 0 0 1rem rgba(255, 212, 121, 0.7);
}

.study-overview {
  display: grid;
  grid-template-columns: minmax(13rem, 0.38fr) minmax(0, 1fr);
  gap: clamp(2rem, 7vw, 6rem);
  padding-block: clamp(4rem, 8vw, 7rem);
  border-bottom: 1px solid rgba(164, 184, 224, 0.1);
}

.study-overview h2 {
  font-size: clamp(1.8rem, 3.8vw, 3rem);
}

.study-description {
  margin: 0;
  color: var(--ink-soft);
  font-size: clamp(1.02rem, 1.8vw, 1.16rem);
  line-height: 1.8;
  white-space: pre-line;
}

.study-description--empty {
  color: var(--ink-muted);
  font-style: italic;
}

.study-map,
.attribution-panel,
.related-section {
  padding-block: clamp(4rem, 8vw, 6.5rem);
  border-bottom: 1px solid rgba(164, 184, 224, 0.1);
}

.study-facts {
  display: grid;
  grid-template-columns: repeat(2, minmax(0, 1fr));
  gap: 1rem;
  margin-top: 2.5rem;
}

.study-fact-card {
  position: relative;
  min-width: 0;
  padding: 1.35rem;
  overflow: hidden;
  border: 1px solid var(--line);
  border-radius: var(--radius-md);
  background: linear-gradient(150deg, rgba(21, 32, 58, 0.84), rgba(9, 15, 30, 0.88));
}

.study-fact-card::after {
  position: absolute;
  top: -3.5rem;
  right: -3.5rem;
  width: 8rem;
  aspect-ratio: 1;
  border: 1px solid rgba(102, 227, 255, 0.1);
  border-radius: 50%;
  content: "";
}

.fact-number {
  margin: 0 0 0.8rem;
  color: var(--cyan);
  font-family: var(--font-mono);
  font-size: 0.67rem;
}

.study-fact-card h3 {
  margin-bottom: 1.1rem;
}

.study-fact-card dl {
  display: grid;
  gap: 1rem;
  margin: 0;
}

.study-fact-card dt,
.attribution-panel dt {
  color: var(--ink-muted);
  font-size: 0.68rem;
  font-weight: 700;
  letter-spacing: 0.06em;
  text-transform: uppercase;
}

.study-fact-card dd {
  margin: 0.35rem 0 0;
  color: var(--ink-soft);
  font-size: 0.84rem;
  overflow-wrap: anywhere;
}

.chip-list {
  display: flex;
  flex-wrap: wrap;
  gap: 0.4rem;
}

.attribution-panel {
  display: grid;
  grid-template-columns: minmax(13rem, 0.38fr) minmax(0, 1fr);
  gap: clamp(2rem, 7vw, 6rem);
}

.attribution-panel h2 {
  font-size: clamp(1.8rem, 3.8vw, 3rem);
}

.attribution-panel dl {
  display: grid;
  gap: 1rem;
  margin: 0;
}

.attribution-panel dl > div {
  padding: 1.1rem;
  border: 1px solid var(--line);
  border-radius: var(--radius-sm);
  background: var(--panel);
}

.attribution-panel dd {
  margin: 0.4rem 0 0;
  color: var(--ink-soft);
  overflow-wrap: anywhere;
}

.related-grid {
  display: grid;
  grid-template-columns: repeat(4, minmax(0, 1fr));
  gap: 0.8rem;
  margin-top: 2.5rem;
}

.related-card {
  display: flex;
  min-height: 12rem;
  flex-direction: column;
  padding: 1.1rem;
  border: 1px solid var(--line);
  border-radius: var(--radius-md);
  background: var(--panel);
  color: var(--ink);
  text-decoration: none;
  transition:
    border-color 160ms ease,
    transform 160ms ease;
}

.related-card:hover {
  border-color: var(--cyan);
  color: var(--ink);
  transform: translateY(-2px);
}

.related-card > span {
  color: var(--cyan);
  font-family: var(--font-mono);
  font-size: 0.65rem;
}

.related-card strong {
  display: -webkit-box;
  margin-top: 0.75rem;
  overflow: hidden;
  font-size: 0.88rem;
  line-height: 1.42;
  -webkit-box-orient: vertical;
  -webkit-line-clamp: 4;
}

.related-card small {
  margin-top: auto;
  padding-top: 1rem;
  color: var(--ink-muted);
  font-size: 0.68rem;
}

.provenance-note {
  margin-top: 3rem;
  padding: 1rem 1.15rem;
  border: 1px solid rgba(169, 149, 255, 0.25);
  border-left: 3px solid var(--violet);
  border-radius: 0 var(--radius-sm) var(--radius-sm) 0;
  background: var(--violet-soft);
  color: var(--ink-soft);
  font-size: 0.84rem;
}

.provenance-note strong {
  color: var(--ink);
}

@keyframes spin {
  to {
    transform: rotate(1turn);
  }
}

/* Responsive -------------------------------------------------------------- */

@media (max-width: 68rem) {
  .metrics-grid,
  .collections-grid {
    grid-template-columns: repeat(2, minmax(0, 1fr));
  }

  .latest-strip {
    grid-template-columns: 1fr;
  }

  .latest-card {
    min-height: auto;
  }

  .explorer-controls {
    grid-template-columns: repeat(2, minmax(0, 1fr));
  }

  .search-field {
    grid-column: 1 / -1;
  }

  .related-grid {
    grid-template-columns: repeat(2, minmax(0, 1fr));
  }
}

@media (max-width: 56rem) {
  .hero,
  .snapshot-panel__header,
  .methodology-section,
  .study-hero,
  .study-overview,
  .attribution-panel {
    grid-template-columns: 1fr;
  }

  .hero-biology {
    display: none;
  }

  .hero {
    min-height: auto;
  }

  .hero::after {
    top: 4rem;
    right: -10rem;
    width: 25rem;
    opacity: 0.5;
  }

  .study-hero__visual {
    min-height: 15rem;
  }

  .study-hero__visual img {
    min-height: 15rem;
    max-height: 25rem;
  }

  .section-heading {
    align-items: flex-start;
    flex-direction: column;
    gap: 0.5rem;
  }
}

@media (max-width: 48rem) {
  html {
    scroll-padding-top: 1rem;
  }

  .site-header {
    position: relative;
  }

  .site-header__inner {
    min-height: auto;
    flex-direction: column;
    align-items: flex-start;
    gap: 0.7rem;
    padding-block: 0.9rem;
  }

  .site-nav {
    width: 100%;
    overflow-x: auto;
    padding-bottom: 0.2rem;
  }

  .site-nav a {
    flex: 0 0 auto;
  }

  h1 {
    font-size: clamp(2.8rem, 14vw, 4.7rem);
  }

  .catalog-layout {
    display: block;
  }

  .filter-toggle {
    display: flex;
    width: calc(100% - 2.5rem);
    min-height: 2.8rem;
    align-items: center;
    justify-content: center;
    margin: 1rem 1.25rem 0;
    padding: 0.55rem 0.9rem;
    border: 1px solid var(--line-strong);
    border-radius: var(--radius-sm);
    background: rgba(164, 184, 224, 0.07);
    color: var(--ink);
    font-size: 0.84rem;
    font-weight: 750;
  }

  .facets {
    position: static;
    display: none;
    max-height: none;
    margin: 1rem 1.25rem 0;
    padding: 1.1rem;
    border: 1px solid var(--line);
    border-radius: var(--radius-md);
    background: #0b1325;
  }

  .facets.is-open {
    display: block;
  }

  .study-grid {
    grid-template-columns: 1fr;
  }

  .study-card {
    min-height: 18rem;
  }

  .study-facts {
    grid-template-columns: 1fr;
  }

  .site-footer__inner {
    grid-template-columns: 1fr;
  }
}

@media (max-width: 38rem) {
  .site-header__inner,
  main > section,
  main > article,
  .site-footer__inner {
    width: min(calc(100% - 1.25rem), var(--content));
  }

  .metrics-grid,
  .collections-grid,
  .explorer-controls,
  .snapshot-stats,
  .methodology-grid,
  .related-grid {
    grid-template-columns: 1fr;
  }

  .snapshot-stats > div {
    border-right: 0;
    border-bottom: 1px solid var(--line);
  }

  .snapshot-stats > div:nth-last-child(-n + 2) {
    border-bottom: 1px solid var(--line);
  }

  .snapshot-stats > div:last-child {
    border-bottom: 0;
  }

  .search-field {
    grid-column: auto;
  }

  .explorer-controls,
  .catalog {
    padding: 0.85rem;
  }

  .active-filters {
    padding-inline: 0.85rem;
  }

  .filter-toggle {
    width: calc(100% - 1.7rem);
    margin-inline: 0.85rem;
  }

  .facets {
    margin-inline: 0.85rem;
  }

  .study-hero {
    min-height: auto;
    padding: 1.25rem;
  }

  .study-hero__visual {
    min-height: 12rem;
  }

  .study-hero__visual img {
    min-height: 12rem;
  }

  .study-hero__meta {
    flex-direction: column;
  }

  .study-hero__meta span + span::before {
    display: none;
  }

  .study-actions {
    align-items: stretch;
    flex-direction: column;
  }

  .study-actions .button {
    width: 100%;
  }

  .pagination button {
    min-width: 2.45rem;
    padding-inline: 0.65rem;
  }
}

@media (prefers-reduced-motion: reduce) {
  *,
  *::before,
  *::after {
    scroll-behavior: auto !important;
    animation-duration: 0.01ms !important;
    animation-iteration-count: 1 !important;
    transition-duration: 0.01ms !important;
  }
}

@media (prefers-contrast: more) {
  :root {
    --ink-soft: #e1e8f8;
    --ink-muted: #bdc8dc;
    --line: rgba(214, 226, 250, 0.42);
    --line-strong: rgba(230, 238, 255, 0.72);
  }
}

@media print {
  :root {
    color-scheme: light;
  }

  body {
    background: #fff;
    color: #111;
  }

  body::before,
  .site-header,
  .explorer-controls,
  .facets,
  .pagination,
  .filter-toggle,
  .active-filters {
    display: none !important;
  }

  main > section,
  main > article {
    width: 100%;
    padding-block: 1.5rem;
  }

  h1,
  h2,
  h3,
  h4,
  .metric__value {
    color: #111;
  }
}